Small Scabious Plume Stenoptilia annadactyla
Apart from the individual below, only known from the Norfolk and Suffolk Brecks where it was first discovered in 2005.
This came to light in a recently established meadow (formerly arable). I thought the overall colour was quite brown and that it would therefore most likely turn out to be Brown Plume Stenoptilia pterodactlya, the only really common species in the genus. But it seemed a bit odd, and there were dark markings in the first lobe that aren't normally present in pterodactyla. Could it be Twin-spot Plume Stenoptilia bipunctidactyla, I mused, and then rejected that idea as it surely wasn't grey enough. I couldn't think of any other alternatives so potted it for subsequent examination at home.
When I came to dissect it I had forgotten it was potentially interesting and thought was just being ultra-cautious checking pterodactyla, and was not quite as careful as I should have been. As a result the genitalia aren't quite so well prepared as I would have liked, but upon seeing the antrum and comparing it with the diagrams in Hart I immediately realised it was not pterodactyla. But it also didn't seem right for bipunctidactyla, and although Dusky Plume Stenoptilia zophodactylus was never really a contender, that was also ruled out leaving only species that I would not have expected to encounter round here, and of those only the apparently rare (or overlooked?) Breckland speciality annadactyla matched.
The antrum shape is key to the identification - the sclerotised portion is noticeably shorter than on the other species (some references seem to refer the sclerotised section as the entire antrum but the membranous section above it seems to me to be part of the same structure so I'm not sure). It is also more parallel-sided, not tapering as it does in the other species.
There also seems to be a difference in the form of the sclerotised line running through the ductus bursae. This isn't referred to in Hart or other references I've seen so maybe it is not diagnostic, but in this individual as well as most of the images of female annadactyla genitalia I've seen, this sclerite is clearly broadened into a more-or-less club shape at its end (the end nearest the corpus bursae). On all the images I've found of Stenoptilia bipunctidactyla it either doesn't broaden at the end at all, or if it does it's not nearly so obvious or abrupt.
Intriguingly, although annadactyla and bipunctidactyla are meant to be more-or-less identical in external appearance, the online images I've seen of annadactyla all show browner (less grey) moths than all the online images I've seen of bipunctidactyla. Of course that's not a large sample, and the photos in Hart show a greyer annadactyla and a (darker) brown bipunctidactyla, so I don't suppose there's much in this if anything, but it will be interesting to see if there's an average difference given a larger sample. Photos of both species show a white-fringed termen with a single dark spot at its inner end - not helpful for separating annadactyla from bipunctidactyla but could help to rule out pterodactyla.
More than 18 miles away from the nearest known Breckland site, this was quite unexpected at this location.
female Small Scabious Plume Stenoptilia annadactyla, Wendling Beck Project (Norfolk, UK), 23rd August 2024
Since finding the above we've established that there are numerous Small Scabious plants growing in the meadow, so I hoped to find more examples of their associated plume moth. None came to light in 2025 but I caught the next two moths together in June 2026. The first one's genitalia pretty much matched the last one:
female Small Scabious Plume Stenoptilia annadactyla, Wendling Beck Project (Norfolk, UK), 23rd June 2026
The second one was slightly fresher - it still had its cilia on the wings for example - and my confidence that it would turn out to be annadactyla based on the wing pattern was higher. But when I saw the genitalia (which were distinctly larger than the other one) I immediately suspected it was in fact something else because the sclerotised line running through the ductus bursae did not widen at the end. There seemed to be other differences too, although in the end I think these were down to differences in the way it was prepared. However after carefully comparing it with images and diagrams/descriptions of genitalia of each of the plausible Stenoptilia species (and some less plausible ones) I concluded that it had to be another annadactyla. The shape of the antrum and the extent that it is sclerotised seems to rule out any other options. In addition to the shape of the antrum and length of sclerotisaton, on this species the sclerotisation fades out before it reaches the junction with the ductus bursae, which doesn't usually seem to be the case (at least to the same extent) on other similar species. Again I'm not sure if that's diagnostic, but it could be a helpful pointer.
female Small Scabious Plume Stenoptilia annadactyla, Wendling Beck Project (Norfolk, UK), 23rd June 2026