Lesser Four-spot Treerunner Calodromius spilotus

The Hackston key says this species is usually found on pine but all of mine have been on Pedunculate Oak.

Lesser Four-spot Treerunner Calodromius spilotus, North Elmham Cathedral Meadows (Norfolk, UK), 21st February 2021

Lesser Four-spot Treerunner Calodromius spilotus, North Elmham Cathedral Meadows (Norfolk, UK), 30th October 2020

Lesser Four-spot Treerunner Calodromius spilotus, North Elmham Cathedral Meadows (Norfolk, UK), 4th January 2020

Lesser Four-spot Treerunner Calodromius spilotus, North Elmham Cathedral Meadows (Norfolk, UK), 9th January 2019 (first 3 photos are the same individual; last photo is a different individual)